diff --git a/webrtc/media/engine/fakewebrtccall.cc b/webrtc/media/engine/fakewebrtccall.cc
index 5b2f0a73c4..c345565b1b 100644
--- a/webrtc/media/engine/fakewebrtccall.cc
+++ b/webrtc/media/engine/fakewebrtccall.cc
@@ -166,9 +166,17 @@ void FakeVideoSendStream::ReconfigureVideoEncoder(
     if (config_.encoder_settings.payload_name == "VP8") {
       vpx_settings_.vp8 = *reinterpret_cast<const webrtc::VideoCodecVP8*>(
                               config.encoder_specific_settings);
+      if (!config.streams.empty()) {
+        vpx_settings_.vp8.numberOfTemporalLayers = static_cast<unsigned char>(
+            config.streams.back().temporal_layer_thresholds_bps.size() + 1);
+      }
     } else if (config_.encoder_settings.payload_name == "VP9") {
       vpx_settings_.vp9 = *reinterpret_cast<const webrtc::VideoCodecVP9*>(
                               config.encoder_specific_settings);
+      if (!config.streams.empty()) {
+        vpx_settings_.vp9.numberOfTemporalLayers = static_cast<unsigned char>(
+            config.streams.back().temporal_layer_thresholds_bps.size() + 1);
+      }
     } else {
       ADD_FAILURE() << "Unsupported encoder payload: "
                     << config_.encoder_settings.payload_name;
diff --git a/webrtc/media/engine/webrtcvideoengine2.cc b/webrtc/media/engine/webrtcvideoengine2.cc
index 395dcb5737..3cff1e6fbd 100644
--- a/webrtc/media/engine/webrtcvideoengine2.cc
+++ b/webrtc/media/engine/webrtcvideoengine2.cc
@@ -10,6 +10,7 @@
 
 #include "webrtc/media/engine/webrtcvideoengine2.h"
 
+#include <stdio.h>
 #include <algorithm>
 #include <set>
 #include <string>
@@ -315,6 +316,44 @@ static int GetMaxDefaultVideoBitrateKbps(int width, int height) {
   }
 }
 
+bool GetVp9LayersFromFieldTrialGroup(int* num_spatial_layers,
+                                     int* num_temporal_layers) {
+  std::string group = webrtc::field_trial::FindFullName("WebRTC-SupportVP9SVC");
+  if (group.empty())
+    return false;
+
+  if (sscanf(group.c_str(), "EnabledByFlag_%dSL%dTL", num_spatial_layers,
+             num_temporal_layers) != 2) {
+    return false;
+  }
+  const int kMaxSpatialLayers = 3;
+  if (*num_spatial_layers > kMaxSpatialLayers || *num_spatial_layers < 1)
+    return false;
+
+  const int kMaxTemporalLayers = 3;
+  if (*num_temporal_layers > kMaxTemporalLayers || *num_temporal_layers < 1)
+    return false;
+
+  return true;
+}
+
+int GetDefaultVp9SpatialLayers() {
+  int num_sl;
+  int num_tl;
+  if (GetVp9LayersFromFieldTrialGroup(&num_sl, &num_tl)) {
+    return num_sl;
+  }
+  return 1;
+}
+
+int GetDefaultVp9TemporalLayers() {
+  int num_sl;
+  int num_tl;
+  if (GetVp9LayersFromFieldTrialGroup(&num_sl, &num_tl)) {
+    return num_tl;
+  }
+  return 1;
+}
 }  // namespace
 
 // Constants defined in webrtc/media/engine/constants.h
@@ -443,6 +482,14 @@ void* WebRtcVideoChannel2::WebRtcVideoSendStream::ConfigureVideoEncoderSettings(
   }
   if (CodecNamesEq(codec.name, kVp9CodecName)) {
     encoder_settings_.vp9 = webrtc::VideoEncoder::GetDefaultVp9Settings();
+    if (is_screencast) {
+      // TODO(asapersson): Set to 2 for now since there is a DCHECK in
+      // VideoSendStream::ReconfigureVideoEncoder.
+      encoder_settings_.vp9.numberOfSpatialLayers = 2;
+    } else {
+      encoder_settings_.vp9.numberOfSpatialLayers =
+          GetDefaultVp9SpatialLayers();
+    }
     // VP9 denoising is disabled by default.
     encoder_settings_.vp9.denoisingOn =
         codec_default_denoising ? false : denoising;
@@ -1908,6 +1955,11 @@ WebRtcVideoChannel2::WebRtcVideoSendStream::CreateVideoEncoderConfig(
     encoder_config.streams[0].temporal_layer_thresholds_bps.push_back(
         config.tl0_bitrate_kbps * 1000);
   }
+  if (CodecNamesEq(codec.name, kVp9CodecName) && !is_screencast &&
+      encoder_config.streams.size() == 1) {
+    encoder_config.streams[0].temporal_layer_thresholds_bps.resize(
+        GetDefaultVp9TemporalLayers() - 1);
+  }
   return encoder_config;
 }
 
diff --git a/webrtc/media/engine/webrtcvideoengine2_unittest.cc b/webrtc/media/engine/webrtcvideoengine2_unittest.cc
index 81f0e5b168..477b62323e 100644
--- a/webrtc/media/engine/webrtcvideoengine2_unittest.cc
+++ b/webrtc/media/engine/webrtcvideoengine2_unittest.cc
@@ -1824,7 +1824,9 @@ TEST_F(WebRtcVideoChannel2Test, VerifyVp8SpecificSettings) {
 
 class Vp9SettingsTest : public WebRtcVideoChannel2Test {
  public:
-  Vp9SettingsTest() : WebRtcVideoChannel2Test() {
+  Vp9SettingsTest() : Vp9SettingsTest("") {}
+  explicit Vp9SettingsTest(const char* field_trials)
+      : WebRtcVideoChannel2Test(field_trials) {
     encoder_factory_.AddSupportedVideoCodecType(webrtc::kVideoCodecVP9, "VP9");
   }
   virtual ~Vp9SettingsTest() {}
@@ -1899,6 +1901,72 @@ TEST_F(Vp9SettingsTest, VerifyVp9SpecificSettings) {
   EXPECT_TRUE(channel_->SetCapturer(last_ssrc_, NULL));
 }
 
+class Vp9SettingsTestWithFieldTrial : public Vp9SettingsTest {
+ public:
+  Vp9SettingsTestWithFieldTrial(const char* field_trials)
+      : Vp9SettingsTest(field_trials) {}
+
+ protected:
+  void VerifySettings(int num_spatial_layers, int num_temporal_layers) {
+    cricket::VideoSendParameters parameters;
+    parameters.codecs.push_back(kVp9Codec);
+    ASSERT_TRUE(channel_->SetSendParameters(parameters));
+
+    FakeVideoSendStream* stream = SetUpSimulcast(false, false);
+
+    cricket::FakeVideoCapturer capturer;
+    EXPECT_EQ(cricket::CS_RUNNING,
+              capturer.Start(capturer.GetSupportedFormats()->front()));
+    EXPECT_TRUE(channel_->SetCapturer(last_ssrc_, &capturer));
+    channel_->SetSend(true);
+
+    EXPECT_TRUE(capturer.CaptureFrame());
+
+    webrtc::VideoCodecVP9 vp9_settings;
+    ASSERT_TRUE(stream->GetVp9Settings(&vp9_settings)) << "No VP9 config set.";
+    EXPECT_EQ(num_spatial_layers, vp9_settings.numberOfSpatialLayers);
+    EXPECT_EQ(num_temporal_layers, vp9_settings.numberOfTemporalLayers);
+
+    EXPECT_TRUE(channel_->SetCapturer(last_ssrc_, NULL));
+  }
+};
+
+class Vp9SettingsTestWithNoFlag : public Vp9SettingsTestWithFieldTrial {
+ public:
+  Vp9SettingsTestWithNoFlag() : Vp9SettingsTestWithFieldTrial("") {}
+};
+
+TEST_F(Vp9SettingsTestWithNoFlag, VerifySettings) {
+  const int kNumSpatialLayers = 1;
+  const int kNumTemporalLayers = 1;
+  VerifySettings(kNumSpatialLayers, kNumTemporalLayers);
+}
+
+class Vp9SettingsTestWithInvalidFlag : public Vp9SettingsTestWithFieldTrial {
+ public:
+  Vp9SettingsTestWithInvalidFlag()
+      : Vp9SettingsTestWithFieldTrial("WebRTC-SupportVP9SVC/Default/") {}
+};
+
+TEST_F(Vp9SettingsTestWithInvalidFlag, VerifySettings) {
+  const int kNumSpatialLayers = 1;
+  const int kNumTemporalLayers = 1;
+  VerifySettings(kNumSpatialLayers, kNumTemporalLayers);
+}
+
+class Vp9SettingsTestWith2SL3TLFlag : public Vp9SettingsTestWithFieldTrial {
+ public:
+  Vp9SettingsTestWith2SL3TLFlag()
+      : Vp9SettingsTestWithFieldTrial(
+            "WebRTC-SupportVP9SVC/EnabledByFlag_2SL3TL/") {}
+};
+
+TEST_F(Vp9SettingsTestWith2SL3TLFlag, VerifySettings) {
+  const int kNumSpatialLayers = 2;
+  const int kNumTemporalLayers = 3;
+  VerifySettings(kNumSpatialLayers, kNumTemporalLayers);
+}
+
 TEST_F(WebRtcVideoChannel2Test, AdaptsOnOveruse) {
   TestCpuAdaptation(true, false);
 }